Cycadaceae cycas family
Stangeriaceae stangeria family
Zamiaceae zamia family
Cycads /ˈsaɪkædz/ are seed plants typically characterized by a stout and woody (ligneous) trunk with a crown of large, hard and stiff, evergreen leaves. They usually have pinnate leaves. The individual plants are either all male or all female (dioecious). Cycads vary in size from having a trunk that is only a few centimeters tall to trunks up to several meters tall. They typically grow very slowly and live very long, with some specimens known to be as much as 1,000 years old. Because of their superficial resemblance, they are sometimes confused with and mistaken for palms or ferns, but are only distantly related to either.
Cycads are found across much of the subtropical and tropical parts of the world. They are found in South and Central America (where the greatest diversity occurs), Mexico, the Antilles, southeastern United States, Australia, Melanesia, Micronesia, Japan, China, Southeast Asia, India, Sri Lanka, Madagascar, and southern and tropical Africa, where at least 65 species occur. Some can survive in harsh semidesert climates (xerophytic), others in wet rain forest conditions, and some in both. Some can grow in sand or even on rock, some in oxygen-poor swampy bog-like soils rich in organic material, and some in both. Some are able to grow in full sun, some in full shade, and some in both. Some are salt tolerant (halophytes).
Cycads belong to the biological division Cycadophyta. There are three extant families of cycads, Cycadaceae, Stangeriaceae, and Zamiaceae. Though they are a minor component of the plant kingdom today, during the Jurassic period they were extremely common. They have changed little since the Jurassic, compared to some major evolutionary changes in other plant divisions.
Cycads are gymnosperms (naked seeded), meaning that their unfertilized seeds (ovules) are open to the air to be directly fertilized by pollination, as contrasted with angiosperms, who have enclosed seeds with more complex fertilization arrangements. Cycads have very specialized pollinators, usually a specific kind of beetle. They have been reported to fix nitrogen in association with a cyanobacterium living in the roots. These blue-green algae produce a neurotoxin called BMAA that is found in the seeds of cycads. A neurotoxin cycling through the food chain from the cyanobacterium through cycads to its seeds, to bats eating the seeds, to humans consuming the bats, is hypothesized to be a source for some neurological diseases in humans.